国家自然基金青年科学基金项目31200320;甘肃农业大学盛彤笙科技创新基金项目GSAU-STS-1332资助 This work was supported by the National Natural Foundation for Young Scientists of China31200320;the Science and Technology Innovation Foundation of Sheng Tongsheng in Gansu Agricultural UniversityGSAU-STS-1332