国家自然科学基金项目31660516;贵州省农业攻关项目[2015]3014-1;贵州省普通高等学校系统与应用蜱螨学创新团队[2014]33;昆虫进化生物学和害虫控制国际科技合作基地[2016]5802;the National Natural Science Foundation of China31660516;Agriculture Research Project of Guizhou Province,China[2015] 3014-1;Innovation Team Program for Systematic and Applied Acarology[2014] 33;the International Cooperation Base for Insect Evolutionary Biology and Pest Control[2016] 5802