国家自然科学基金项目31600395;云南省教育厅重点基金项目2015Z136;云南省高原湿地科学创新团队项目2012HC007;the National Natural Science Foundation of China31600395;the Key Program of the Educational Department of Yunnan Province2015Z136;the Plateau Wetlands Science Innovation Team of Yunnan Province2012HC007