国家重点研发计划项目2016YFD0201100;国家现代农业产业技术体系建设资金项目CARS-28;国家自然科学基金项目31501713;山东农业大学博上后基金项目010-76513资助 This work was supported by the Key Research and Development Program of China2016YFD0201100;the Earmarked Fund for the China Agriculture Research SystemCARS-28;the National Natural Science Foundation of China31501713;the Postdoctoral Science Foundation of Shandong Agricultural University010-76513