海洋公益性行业科研专项201105009-3;国家科技支撑项目2011BAD13B08;浙江省新苗人才项目2014R405047;宁波大学水产养殖浙江省重中之重一级学科开放基金项目资助421500052 This work was supported by the Science and Technology Research Project of Ocean in the Public Interest201105009-3;the National Science & Technology Pillar Program2011BAD13B08;the Zhejiang Province Xinmiao Talents Project2014R405047;the Zhejiang Open Foundation of the Most Important Subjects for Aquaculture in Ningbo University421500052