本文由中央级公益性科研院所基本科研业务费专项CAFYBB2014MA004和国家重点研发计划项目2016YFC0502104-02资助 This work was supported by the National Nonprofit Institute Research Grant of Chinese Academy of Forestry CAFYBB2014MA004,and the National Key Research and Development Program 2016YFC0502104-02.