本文由国家科技支撑计划项目2013BAD13B03、公益性行业农业科研专项201203018、国家自然基金项目41606210和教育部留学回国人员科研启动基金项目资助;This paper was supported by the National Science & Technology Support Plan of China 2013BAD13B03,Special Fund for Argo-scientific Research in the Public Interest of China 201203018,National Natural Science Foundation of China 41606210 and Scientific Research Foundation for the Returned Overseas Chinese Scholars of Ministry of Education.