本文由安徽省烟草公司科技项目20140551004、安徽省农业科学院院长青年基金项目14B0945和公益性行业农业科研专项201203013资助;This paper was supported by Anhui Tobacco Corporation 20140551004,2014 Dean Younth Innovation Fund of Anhui Academy of Agricultural Sciences 12B0919 and the National Department Public Benefit Research Foundation of China 201203013.