本文由国家自然科学基金项目81303170,U1205021,31401950,81573530、福建农林大学杰出青年科研人才计划项目XJQ201501和福建省省级扶贫重点县科技人员专项计划项目K1515118A资助,This work was supported by the National Natural Science Foundation of China 81303170,U1205021,31401950,81573530,the Outstanding Youth Scientific Fund of Fujian Agriculture and Forestry UniversityXJQ201501 and the Science and Technology Project for Fujian Poverty Alleviation K1515118A.