本文由现代农业产业技术体系建设专项CARS-03-2-7和国家科技支撑计划项目2015BAD22B03-03资助 This work supported by the Project of Technology System in Modern Wheat Industry,Ministry of Agriculture,China CARS-03-2-7 and the National Science & Technology Support Program of China 2015BAD22B03-03.