本文由国家“948”引进国际先进林业科学技术计划项目2011-4-60和山东省农业重大应用技术创新计划项目鲁财农指[2016]36号资助;This work was supported by the National 948 Program of the Introduction of International Advanced Forestry Technology,China 2011-4-60 and the Agricultural Major Application Technology Innovation Program of Shandong Province Financial and Agricultural Indicators,2016,No.36.