本文由国家自然科学基金项目31400406、海洋公益性行业科研专项201405010和国家重点基础研究发展计划项目2011CB403601资助 This work was supported by the National Natural Science Foundation of China 31400406,the National Ocean Welfare Research 201405010 and the National Key Basic Research and Development Plan 2011CB403601.