本文由国家自然科学基金项目31300405、广东省科技计划项目2015B090903077、教育部人文社科研究项目13YJCZH015和广东省现代农业产业技术体系建设专项粤农[2009]380号资助 This work was supported by the National Natural Science Foundation of China 31300405,the Science and Technology Program of Guangdong Province 2015B090903077,the Humanities and Social Sciences Research Project of the Ministry of Education of China 13YJCZH015,and the Special Subject of Construction of National System of Modern Industrial Technology of Guangdong Province Guangdong Agriculture [2009] 380.