本文由国家科技支撑计划项目2015BAD22B02、公益性行业农业科研专项201303104,201503116和国家自然科学基金项目31571620资助 This work was supported by National Science and Technology Support Program 2015BAD22B02,the Special Fund for Agro-scientific Research in the Public Interest 201303104,201503116,and the National Natural Science Foundation of China 31571620.