本文由公益性行业农业科研专项201503125-02、中澳国际合作项目SMCN/2002/094和甘肃省农业科技创新项目GNCX-2014-35资助 This work was supported by the Agro-scientific Research Programs in Public Interest 201503125-02,the China-Australia for International Cooperation Program SMCN/2002/094,and the Innovation Program of the Agricultural Science and Technology of Gansu Province,China GNCX-2014-35.