本文由国家重点基础研究发展计划项目2011CB403601、国家自然科学基金项目31400406,31302173、辽宁省自然科学基金项目2015020795和国家农业科技成果转化资金项目2013GB2B000096资助 This work was supported by the National Key Basic Research and Development Plan 2011CB403601,National Natural Science Foundation of China 31400406,31302173,the Natural Science Foundation of Liaoning Pro-vince 2015020795,and the National Agricultural Scientific Achievement Transfer Fund 2013GB2B000096.