本文由国家“十二五”科技支撑计划项目2014BAD05B03、山东省农业重大应用技术创新项目2013-136和山东省优秀中青年科学家科研奖励基金项目BS2014NY005资助;This work was supported by 12th Five-Year State Science and Technology Support Program 2014BAD05B03,the Key Technology Innovation Project for Agriculture of Shandong Province 2013-136 and the Excellent Young Scientist Foundation of Shandong Province BS2014NY005.