本文由国家自然科学基金项目41371508、陕西省自然科学基础研究计划重点项目2013JZ006和西北农林科技大学基本科研业务费专项ZD2013021资助;This paper was supported by the National Nature Science Foundation of China 41371508,the Nature Science Basic Research Key Project of Shaanxi Province,China 2013JZ006 and the Special Funds of the Basic Scientific Research Project of NWSUAF,China ZD2013021.