本文由国家自然科学基金项目31401334、农业部现代小麦产业技术体系项目CARS-3-1-19和山东省科技发展计划项目2014GNC111017资助 This work was supported by the National Natural Science Foundation of China 31401334,the Project of Technology System in Modem Wheat Industry,Ministry of Agriculture,China CARS-3-1-19 and the Project of Science and Technology Development of Shandong Province 2014GNC111017.