本文由河北省应用基础研究计划重点项目13963302D,15963301D、河北省创新能力提升计划项目154576235和海洋公益性行业科研专项201005009资助 This work was supported by the Key Basic Research Program of Hebei Province13963302D,15963301D,the Innovation Ability Promotion Project of Hebei Province 154576235 and the State Oceanic Administration of Social Public Welfare Project 201005009.