本文由国家自然科学基金项目31170487、中央高校基本科研业务费项目KYZ201207和南京农业大学SRT项目1413A09资助 This work was supported by the National Natural Science Foundation of China 31170487,the Central Universities Fundamental Research Project KYZ201207 and SRT Project of Nanjing Agricultural University 1413A09.