国家自然科学基金编号:11371333;山东省自然科学基金编号:ZR2013FQ026;中央高校基本科研业务费专项编号:201362033;海洋公益性行业科研专项经费项目编号:201505002 Supported by National Natural Science Foundation of China No.11371333;Shandong Provincial Natural Science Foundation,China No.ZR2013FQ026;Public Science and Technology Research Funds Projects of Ocean No.201505002