Natural Science Foundation of China31371577;Natural Science Foundation of Youth Fund of Shandong ProvinceZR2014CQ040;Science and Technology Development Plan of Tai'an201440774-11B;Agricultural Seed Project of Tai'an No.2015LZ06.国家自然科学基金31371577;山东省自然科学基金青年基金ZR2014CQ040;泰安市科技发展计划201440774-11B;泰安市农业良种工程2015LZ06