国家自然基金青年基金71303038;国家自然基金国际合作交流项目7141101042;黑龙江省普通本科高校青年创新人才培养计划;东北农业大学“学术骨干”项目.The Natural Foundation Youth Fund Project71303038;the National Natural Science Foundation of International Cooperation and Exchange7141101042;the Heilongjiang Province Undergraduate Youth Innovative Talents Training Program;the Northeast Agricultural University "Academic Backbone" Project