国家自然科学基金31401349;陕西省科技统筹创新工程计划2015KTZDNY01-02;陕西省科学技术研究发展计划农业攻关项目2014K01-02-01;西北农林科技大学基本科研业务费专项2014YB078.National Natural Science Foundation of China31401349;Shaanxi Province Overall-plan Innovation Project of Science and Technology No.2015KTZDNY01-02,Science and Technology Research and Development Project of Shaanxi Province2014K01-02-01;Northwest Agriculture and Forestry University Basic Scientific Research Business Expenses Special Funds Project2014YB078