National Key Research and Development Program of China2016YFD0500101;National Natural Science Foundation of China31402219,315725199;Shanghai Key Project on Agricultural Development No.2015-6-1-9国家重点研发计划项目2016YFD0500101;国家自然科学基金项目31402219,315725199;上海市科技兴农重点攻关项目沪农科攻字2015第6-1-9号