国家自然科学基金31400473;山东省林业科技创新计划LYCX07-2018-36;山东省高等学校科技计划J14LE16Supported by the National Natural Science Fundation of China31400473;by the Forestry Science and Technology Innovation Plan of Shandong ProvinceLYCX07-2018-36;by the Science and Technology Plan Project for Colleges and Universities in Shandong Province of ChinaJ14LE16