the Youth Science Foundation of Heilongjiang ProvinceQC2014C020;by the Science and Technology Project of Mudanjiang CityZ2016n0015;by the School Cultivation Subject of Heilongjiang Bayi Agricultural University XZR2016-15黑龙江省青年科学基金QC2014C020;牡丹江市科学技术计划项目Z2016n0015;黑龙江八一农垦大学“校内培育课题资助计划”项目XZR2016-15