国家自然科学基金31502074;教育部“促进与美大地区科研合作与高层次人才培养项目”教外司美[2014]2029号;公益性行业农业科研专项201303033;贵州大学引进人才科研项目贵大人基合字201410号Supported by the National Natural Science Foundation of China 31502074,by the Scientific Research Cooperation Project with America and Oceania Region,Ministry of Education JWSM-2014-2029,by the Chinese Special Fund for Agro-scientific Research in the Public Interest 201303033 and by the Scientific Research Project of Guizhou University Talents Fund GDRJHZ-2014-10