国家自然科学基金31300082,31470244;浙江农林大学科研发展基金2013FR012;浙江省2015年度高校国内访问学者专业发展项目178Supported by National Natural Science Foundation of China 31300082,31470244,by the Science Development Foundation of Zhejiang Agricultural and Forestry University 2013FR012 and by the National Visiting Scholar Development Program of Zhejiang 2015 #178