国家自然科学基金31160006;云南省科技条件平台建设计划项目2009DA002;国家微生物资源平台专项-面向微生物教学实验的专题服务项目NIMR-2014-8Supported by the National Natural Science Foundation of China 31160006,the Yunnan Provincial Sciences and Technology Department 2009DA002,National Infrastructure of Natural Resources for Science and Technology Program of China NIMR-2014-8