Supported by the Key Project of Chinese National Programs for Fundamental Research and Development 2011CB111607,by the Natural Science Foundation of China 31370059,by the Special Fund for Agro-scientific Research in the Public Interest 201303094 and by the International Cooperation Project of Chinese Ministry of Science and Technology 2012DFA31120;科技部973前期项目2011CB111607;国家自然科学基金项目31370059;公益性行业农业科研专项201303094;科技部国际合作项目2012DFA31120