山东省优秀中青年科学家科研奖励基金BS2011NJ018;国家自然科学基金40801193;中国海洋大学本科生研究发展计划1312011006;Supported by Scientific Research Foundation for the Excellent Middle-Aged and Youth Scientists of Shandong Province of ChinaBS2011NJ018,National Natural Science Foundation of China 40801193 and by the Undergraduate Research Program of Ocean University of China 1312011006