科技部国际合作专项项目“大气组成变化及其影响与对策研究”2010DFA22770;中国科学院应用微生物研究网络项目KSCX2-EW-G-16;Supported by the International Science Cooperation and Communication Program of China 2010DFA22770 and by the Research Network for Applied Microbiology of the Chinese Academy of Sciences KSCX2-EW-G-16