国家自然科学基金31270143;黑龙江大学高层次人才创新团队支持计划Hdtd2010-17;黑龙江省教育厅科学技术研究项目12511423;黑龙江省高等学校科技创新团队;the National Natural Science Foundation of China31270143;by the High-level Talents innovation team Projects of Heilongjiang UniversityHdtd2010-17;by the Education Department of Heilongjiang Province Scientific Technology Research Project12511423;by the Scientific and Technological Innovation Team Agricultural Microbe Fermentation Technologyof Colleges and Universities of Heilongjiang Province