the Special Fund for Agro-scientific Research in the Public Interest of China201203085;by the Key Program of Science and Technology of Zhejiang Province of China2012C12009-4;by the Natural Science Foundation of Huzhou 2013YZ02农业部公益性行业科研专项201203085;浙江省科技厅重大科技项目2012C12009-4;湖州市自然科学基金2013YZ02