the National Key Project of Fundamental Scientific Research2011CB111607;by the Program of International Science & Technology Cooperation the International Cooperation Project2012DFA31120;by the National "948" Project2011-G25;by the National Natural Science Foundation of China 31370059国家“973项目”前期2011CBl11607;科技部国际合作基金2012DFA31120;农业部“948项目”2011-G5;国家自然科学基金31370059