广东省自然科学基金项目2015A030308015;广东省野生动植物保护管理项目;广东省省级科技计划项目2015A020220011。This work was supported by grants from the Guangdong Natural Science Foundation2015A030308015;Guangdong Wildlife Conservation and Management Project,and Science and Technology Planning Project of Guangdong Province2015A020220011