国家重点研发计划项目2017YFD0201801;吉林省科技厅重大科技招标专项20150203004NY;吉林农业大学博士启动基金项目201634资助 Supported by the National Basic Research Program of China2017YFD0201801;the Special Science and Technology Bidding for Science in Jilin Province20150203004NY;the Doctoral Fund of Jilin Agriculture University201634