国家科技支撑计划项目2013BAD07B11;公益性行业农业科研专项经费项目201203013-06;湖南省自然科学基金项目2016JJ6063资助 Supported by the National Key Technology R&D Program of China2013BAD07B11;the Special Fund for Agro-scientific Research in the Public Interest of China201203013-06;the Natural Science Foundation of Hunan Province of China2016JJ6063