国家自然科学基金项目41471225;辽宁省农业领域青年科技创新人才培养计划2014054;辽宁省高等学校优秀人才支持计划LJQ2013074共同资助 Supported by the National Natural Science Foundation of China41471225;the Cultivation Plan for Youth Agricultural Science and Technology Innovative Talents of Liaoning Province2014054;the Talents Support Program of Higher Learning Institutions in Liaoning Province of ChinaLJQ2013074