国家自然科学基金项目41671321;国家重点研发计划项目2016YFD0800304;山东农业大学青年创新基金23816资助 Supported by the National Natural Science Foundation of China41671321;the National Key Research and Development Project of China2016YFD0800304;the Science and Technology Innovation Foundation for Youth of Shandong Agricultural University23816