中国科学院前沿科学重点研究项目QYZDY-SSW-SMC025;现代农业产业技术体系NYCYTX-49;福建省自然科学基金面上项目2016J01161资助 [Supported by Key Research Program of Frontier Sciences, CASQYZDY-SSW-SMC025;the Special Fund of China Agriculture Research SystemNYCYTX-49;the Natural Science Foundation of Fujian Province, China 2016J01161]