十二五国家科技计划农村领域项目2012BAD25B04;公益性行业农业科研专项201203083;农业部热带亚热带水产资源利用与养殖重点实验室开放课题资助 Supported by the National Great Project of Scientific and Technical Supporting Program2012BAD25B04;Special Fund for Agro Scientific Research in the Public Interest201203083;Open Topic of Key Laboratory of Tropical and Subtropical Fishery Resource Application and Cultivation, Ministry of Agriculture