国家自然科学基金31402329、31174238和U1205123;福建省自然科学基金2014J05042、2012J06008和JK2014026;鳗鲡工程中心科研基金资助 Supported by the National Natural Science Foundation of China31402329, 31174238 and U1205123;the Natural Science Foundation of Fujian Province2014J05042, 2012J06008 and JK2014026;a Fund from Engineering Research Center of the Modern Technology for Eel Industry