现代农业产业技术体系建设专项资金CARS-46-08;国家自然科学基金31272695;湖北省重大科技创新计划项目2013ABA004资助[Supported by Special Funds For the Construction of Modem Agricultural Industry Technology SystemCARS-46-08;the National Natural Science Foundation of China31272695;Major Scientific and Technological Innovation Projects in Hubei Province 2013ABA004]