上海市科学技术委员会科研项目13320502100,13231203504;国家863高技术研究发展计划项目2012AA10A409;科技部港澳台科技合作专项项目2014DFT30270;上海高校水产学一流学科建设项目2012-62-0908资助[Supported by Research Project of Shanghai Scientific and Technical CommitteeGrant 13320502100,13231203504;National High-tech Research and Development Program of China 863 ProgramGrant 2012AA10A409;Hong Kong,Macao and Taiwan Science and Technology Cooperation Program of the Ministry of Science and TechnologyGrant 2014DFT30270;Project of Shanghai Universities First-class Disciplines Project of Fisheries Grant No.2012-62-0908]