中国水产科学研究院基本科研业务费资助项目2015C07XK01;国家科技支撑计划项目2015BAD17B02Supported by Special Scientific Research Funds for Central Non-profit Institutes,Chinese Academy of Fishery Sciences2015C07XK01;the National Key Technology Research and Development Program of the Ministry of Science and Technology of China2015BAD17B02