国家自然科学基金重大国际合作项目31210103918;国家自然科学基金重点项目31430088;湖南省生物发育工程及新产品研发协同创新中心20134486;长沙市科技计划项目K1205009-21 National Natural Science Foundation of China31430088 and 31210103918;Cooperative Xnnovation Center of Engineering and New Products for Developmental Biology20134486;Science-Technology Foundation of Changsha, Hunan Province, ChinaK1205009-21