国家“八六三”高技术研究发展计划2012AA10A411;国家自然科学基金31072208;国家农业科技成果转化资金2013GB2C220537;福建省省长专项基金2014S1477-10National High Technology Research & Development Program of China “863”Program2012AA10A411;National Natural Science Foundation of China31072208;National Agricultural Science and Technology Achievements Transformation Project2013GB2C220537;Special Fund of Fujian Provincial Governor2014S1477-10